Once in the mission press Esc to get the menu, click on the "Start recording" label, a windows pop-up will appear asking you where to save the track file and how to name it (just like saving a text file in notepad).
Give it a name and a saving location, press ok and return to the sim.
I'm not 100% sure but i think that in order for this to work you need to have the windows aero interface enabled (otherwise the windows pop-up might crash). It was the same when selecting aircraft in multiplayer untill a few patches ago, then they made an in-game interface for that and don't use windows pop-ups anymore, so maybe they'll make something similar for the track recorder too.
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